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Spies, J. J., S. M. C. v. Wyk, I. C. Nieman & E. J. L. Liebenberg. 1997. Cytogenetic studies in some representatives of the subfamily Pooideae (Poaceae) in South Africa. 3. The tribe Poeae. Bothalia 27: 75–82.
Spies, J. J., T. H. Burger & S. M. C. v. Wyk. 1999. Chromosome studies on African plants. 12. The tribes of subfamily Pooideae. Bothalia 29(2): 335–341.